Indeed, I was unable to find any evidence whatsoever of the occurrence of a drastic evolutionary acceleration and genetic reconstruction in widespread, populous species.

The major novelty of my theory was its claim that the most rapid evolutionary change does not occur in widespread, populous species, as claimed by Most geneticists, but in small founder populations.

Most of them are doomed to rapid extinction, but a few may make evolutionary inventions, such as physiological, ecological, or behavioral innovations that give these species improved competitive potential.

All I claimed was that when a drastic change occurs, it occurs in a relatively small and isolated population.

I had found again and again that the most aberrant population of a species - often having reached species rank, and occasionally classified even as a separate genus - occurred at a peripheral location, indeed usually at the most isolated peripheral location.

It seems to me that for Darwin the pulsing of evolutionary rates was a strictly vertical phenomenon.